My research integrates genomic data with experiments employing ecological, behavioral, and physiological methods to establish links between genotype, phenotype, and fitness in natural systems. Much of my research has centered on two major questions: (1) how do particular ecological factors influence adaptation? and (2) what controls the repeatability of adaptation across levels of biological organization? To answer these questions I study a variety of systems, often focusing on replicated populations adapting to clearly defined ecological regimes in natural or semi-natural conditions.